Date of Award

Summer 8-15-2025

Level of Access Assigned by Author

Open-Access Thesis

Degree Name

Doctor of Philosophy (PhD)

Department

Biochemistry and Molecular Biology

First Committee Advisor

Guangwen Ren

Second Committee Member

Lenny Shultz

Third Committee Member

Chih-Hao Chang

Additional Committee Members

Sheng Li

Xia Liu

Abstract

Aging is a major risk factor for breast cancer metastasis, yet the mechanisms by which it alters distant organ microenvironments remain poorly understood. Here, we used murine models of pre-metastatic lung niche to investigate how aging reprograms stromal and immune compartments to promote pulmonary metastasis.

Single-cell RNA sequencing revealed that fibroblasts in aged lungs acquire immunoregulatory functions, including upregulation of antigen presentation genes and interferon-stimulated pathways, but lack costimulatory signals, potentially inducing T cell anergy or Treg expansion.

Aged fibroblasts also showed elevated prostaglandin E2 (PGE2) production, which may impair dendritic cell activation and promote suppressive neutrophil phenotypes in co-culture. In parallel, myeloid cells exhibited functional shifts: alveolar macrophages showed ECM remodeling signatures, dendritic cells had reduced antigen-presenting capacity, and neutrophils displayed enhanced NETosis and immunecheckpoint expression. Together, these alterations formed a self-reinforcing loop of stromal–myeloid crosstalk that established an immunosuppressive, pro-metastatic niche.

Our results underscore the importance of aging fibroblasts in shaping an immunosuppressive lung environment and provide potential targets such as the PGE2–EP axis for therapeutic intervention to mitigate metastatic risk in elderly cancer patients.

Available for download on Saturday, September 19, 2026

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